Derek and Casey are back and continue their hilarious sibling rivalry during a family vacation to visit their grandmother at her beautiful lake front lodge. Things heat up with Casey when she meets Jesse, a handsome young dancer who is a waiter at the lodge. Meanwhile as Derek is "chilling" on his vacation, he falls for Roxy, the rich girl who lives across the lake. But when Roxy's father, a greedy land developer, threatens to destroy the natural beauty around them, the McDonald-Venturi kids band together to save the lake and the lodge.
Anne, now a middle-aged woman, is troubled by recent events in her life. When a long-hidden secret is discovered under the floorboards at Green Gables, Anne retreats into her memories to relive her troubled early years prior to arriving as an orphan at Green Gables and being adopted by the Cuthberts.